Koos Biesmeijer
Professor Natural Capital
- Name
- Prof.dr. J.C. Biesmeijer
- Telephone
- +31 71 527 2727
- j.c.biesmeijer@cml.leidenuniv.nl
- ORCID iD
- 0000-0003-0328-0573
Professor Natural Capital
News
Former PhD Candidates
Professor Natural Capital
- Science
- Centrum voor Milieuwetenschappen Leiden
- CML/Environmental Biology
- Pan K., Marshall L., Snoo G.R. de & Biesmeijer J.C. (2024), Dutch landscapes have lost insect-pollinated plants over the past 87 years, Journal of Applied Ecology 61(6): 1323-1333.
- Wouters R.M., Beukema W., Schrama M.J.J., Biesmeijer J.C., Braks M.A.H., Helleman P.J.J., Schaffner F. Slobbe J. van Stroo A. & Beek J.G. van der (2024), Local environmental factors drive distributions of ecologically-contrasting mosquito species (Diptera: Culicidae), Scientific Reports 14: 19315.
- Moens M., Biesmeijer J.C., Klumpers S.G.T. & Marshall L. (2023), Are threatened species special? : An assessment of Dutch bees in relation to land use and climate, Ecology and Evolution 13(7): e10326.
- Pan K., Marshall L., Biesmeijer J.C. & Snoo G.R. de (2022), The distributions of insect, wind and self pollination of plants in the Netherlands in relation to habitat types and 3D vegetation structure, Journal of Pollination Ecology 31(3): 16-28.
- Gogaladze A., Son M.O., Lattuada M., Anistratenko V.V., Syomin V.L., Pavel A.B., Popa O.P., Popa L.O., Poorten J.J. ter, Biesmeijer J.C., Raes N., Wilke T., Sands A.F., Trichkova T., Hubenov Z.K., Vinarski M.V., Anistratenko O.Y., Alexenko T.L. & Wesselingh F.P. (2021), Decline of unique Pontocaspian biodiversity in the Black Sea Basin: A review, Ecology and Evolution 11(19): 12923-12947.
- Biesmeijer J.C. & Bodegom P.M. van (6 January 2021), Exploring Natural Capital Accounting with Koos Biesmeijer and Peter van Bodegom Interviewed by [interview].
- Pan K., Moens M., Marshall L., Cieraad E., Biesmeijer K. & Snoo G.R. de (2021), Importance of natural land cover for plant species’ conservation: a nationwide study in The Netherlands, PLoS ONE 16(11): e0259255.
- Biesmeijer J.C., Moens M., Zelfde M. van 't. & Wit F. (2020), De nulsituatie van biodiversiteit in het werkgebied van Hoogheemraadschap van Rijnland: inzichten en handelingsperspectief ter verbetering van biodiversiteit op basis van monitoring van bestuivers en vegetatie in AWZI-terreinen en analyse van waterplanten no. 2020-021. Leiden: Pollinator Ecology Group, Naturalis BC.
- Cole L.J., Kleijn D., Dicks L.V., Stout J.C., Potts S.G., Albrecht M., Balzan M.V., Bartomeus I., Bebeli P.J., Bevk D., Biesmeijer J.C., Chlebo R., Dautarte A., Emmanouil N., Hartfield C., Holland J.M., Holzschuh A., Knoben N.T.J., Kovacs-Hostyanszki A., Mandelik Y., Panou H., Paxton R.J. Petanidou T., de Carvalho M.A.A.P., Rundlof M., Sarthou J.P., Stavrinides M.C., Suso M.J., Szentgyorgyi H., Vaissiere B.E., Varnava A., Vila M., Zemeckis R. & Scheper J. (2020), A critical analysis of the potential for EU Common Agricultural Policy measures to support wild pollinators on farmland, Journal of Applied Ecology 57: 681-694.
- Marshall L., Perdijk F., Dendoncker N., Kunin W., Roberts S. & Biesmeijer J.C. (2020), Bumblebees moving up: shifts in elevation ranges in the Pyrenees over 115 years, Proceedings of the Royal Society B: Biological Sciences 287(1938): 20202201.
- Ferreira P.A., Boscolo D., Lopes L.E., Carvalheiro L.G., Biesmeijer J.C., da Rocha P.L.B. & Viana B.F. (2020), Forest and connectivity loss simplify tropical pollination networks, Oecologia 192(2): 577-590.
- Tanis M.F., Marshall L., Biesmeijer J.C. & Kolfschoten L. van (2020), Grassland management for meadow birds in the Netherlands is unfavourable to pollinators, Basic and Applied Ecology 43: 52-63.
- Carvalheiro L.G., Biesmeijer J.C., Franzén M., Aguirre-Gutiérrez J., Garibaldi L.A., Helm A., Michez D., Pöyry J., Reemer M., Schweiger O., Berg L. van den, Wallis de Vries M.F. & Kunin W.E. (2020), Soil eutrophication shaped the composition of pollinator assemblages during the past century, Ecography 43(2): 209-221.
- Gogaladze A., Raes N., Biesmeijer J.C., Ionescu C., Pavel A.-B., Son M.O., Gozak N., Anistratenko V.V. & Wesselingh F.P. (2020), Social network analysis and the implications for Pontocaspian biodiversity conservation in Romania and Ukraine: a comparative study, PLoS ONE 15(10): e0221833.
- Gogaladze A., Wesselingh F.P., Biesmeijer K., Anistratenko V.V., Gozak N., Son M.O. & Raes N. (2020), Using social network analysis to assess the Pontocaspian biodiversity conservation capacity in Ukraine, Ecology & Society 25(2): 25.
- Turnhout E., Oudenhoven A. van, Willemen L., Visseren-Hamakers I. & Biesmeijer K. (18 May 2019), Ook u kunt bijdragen aan natuurbehoud. Trouw: 27.
- Cieraad E., Zelfde M. van 't & Biesmeijer J.C. (2019), Biodiversiteitsonderzoek in het kader van de omgevingsvisie in Hart van Holland. CML reports no. 196. Leiden: Institute of Environmental Sciences (UL-CML).
- Yasrebi‐de Kom I.A.R., Biesmeijer J.C. & Aguirre‐Gutiérrez J. (2019), Risk of potential pesticide use to honeybee and bumblebee survival and distribution: A country‐wide analysis for The Netherlands, Diversity and Distributions 25(11): 1709-1720.
- Aizen M.A., Aguiar S., Biesmeijer J.C., Garibaldi L.A., Inouye D.W., Jung C., Martins D.J., Medel R., Morales C.L., Ngo H., Pauw. A., Paxton R.J., Sáez A. & Seymour C.L. (2019), Global agricultural productivity is threatened by increasing pollinator dependence without a parallel increase in crop diversification, Global Change Biology 25(10): 3516-3527.
- Movalli P., Duke G., Ramello G., Dekker R., Vrezec A., Shore R.F., Garcia-Fernandez A., Wernham C., Krone O., Alygizakis N., Badry A., Barbagli F., Biesmeijer J.C., Boano G., Bond A.L., Choresh Y., Christensen J.B., Cincinelli A., Danielsson S., Dias A., Dietz R., Eens M., Espin S., Eulaers I., Frahnert S., Fuiz T.I., Gkotsis G., Glowacka N., Gomez-Ramirez P., Grotti M., Guiraud M., Hosner P., Johansson U., Jaspers V.L.B., Kamminga P., Koschorreck J., Knopf B., Kubin E., LoBrutto S., Lourenco R., Martellini T., Martinez-Lopez E., Mateo R., Nika M.C., Nikolopoulou V., Osborn D., Pauwels O., Pavia M., Pereira M.G., Rudel H., Sanchez-Virosta P., Slobodnik J., Sonne C., Thomaidis N., Topfer T., Treu G., Vainola R., Valkama J., Mije S. van der, Vangeluwe D., Warren B. & Woog F. (2019), Progress on bringing together raptor collections in Europe for contaminant research and monitoring in relation to chemicals regulation, Environmental Science and Pollution Research 26: 20132-20136.
- Kleijn D., Biesmeijer J.C., Dupont Y.L., Nielsen A., Potts S.G. & Settele J. (2018), Bee conservation: inclusive solutions, Science 360(6387): 389-390.
- Kleijn D., Linders T.E.W., Stip A., Biesmeijer J.C., Wackers F.L. & Bukovinszky T. (2018), Scaling up effects of measures mitigating pollinator loss from local- to landscape-level population responses, Methods in Ecology and Evolution 9(7): 1727-1738.
- Marshall L., Biesmeijer J.C., Rasmont P., Vereecken N.J., Dvorak L., Fitzpatrick U., Francis F., Neumayer J., Odegaard F., Paukkunen J.P.T., Pawlikowski T., Reemer M., Roberts S.P.M., Straka J., Vray S. & Dendoncker N. (2018), The interplay of climate and land use change affects the distribution of EU bumblebees, Global Change Biology 24(1): 101-116.
- Biesmeijer J.C., Aquirre-Guttierez J., Zelfde M. van 't, Knoben A., Steen S. van der & Groot A. de (2018), Final Report Honeybee Surveillance Program the Netherlands 2014-2018. Leiden: Naturalis BC.
- Bukovinszky T., Rikken I., Evers S., Wackers F.L., Biesmeijer J.C., Prins H.H.T. & Kleijn D. (2017), Effects of pollen species composition on the foraging behaviour and offspring performance of the mason bee Osmia bicornis (L.), Basic and Applied Ecology 18: 21-30.
- Bukovinszky T., Verheijen J., Zwerver S., Klop E., Biesmeijer J.C., Wackers F.L., Prins H.H.T. & Kleijn D. (2017), Exploring the relationships between landscape complexity, wild bee species richness and reproduction, and pollination services along a complexity gradient in the Netherlands, Biological Conservation 214: 312-319.
- Giannini T.C., Costa W.F., Cordeiro G.D., Imperatriz-Fonseca V.L., Saraiva A.M., Biesmeijer J. & Garibaldi L.A. (2017), Projected climate change threatens pollinators and crop production in Brazil, PLoS ONE 12: e0182274.
- Gillespie M.A.K., Baude M., Biesmeijer J., Boatman N., Budge G.E., Crowe A., Memmott J., Morton R.D., Pietravalle S., Potts S.G., Senapathi D., Smart S.M. & Kunin W.E. (2017), A method for the objective selection of landscape-scale study regions and sites at the national level, Methods in Ecology and Evolution 8(11): 1468-1476.
- Aguirre-Gutierrez J., Kissling W.D., Biesmeijer J.C., WallisDeVries M.F., Reemer M. & Carvalheiro L.G. (2017), Historical changes in the importance of climate and land use as determinants of Dutch pollinator distributions, Journal of Biogeography 44(3): 696-707.
- Aquirre-Gutierrez J., Wallis de Vries M.F., Marshall L., Zelfde M. van 't, Villabos-Arambula A.R., Boekelo B., Bartholomeus H., Franzen M. & Biesmeijer J.C. (2017), Butterflies show different functional and species diversity in relationship to vegetation structure and land use, Global Ecology and Biogeography 26(10): 1126–1137.
- Groot G.A. de, Knoben N., Kats R. van, Dimmers W., Zelfde M. van 't, Reemer M., Biesmeijer K. & Kleijn D. (2016), De bijdrage van (wilde) bestuivers aan een hoogwaardige teelt van peren en aardbeien. Nieuwe kwantitatieve inzichten in de diensten geleverd door de bestuivende insecten aan de fruitteeltsector in Nederland. Alterra no. 2716. Wageningen: Alterra Wageningen UR.
- Cormont A., Siepel H., Clement J., Melman T.C.P., Wallis de Vries M.F., Turnhout C.A.M. van, Sparrius L.B., Reemer M., Biesmeijer J.C., Berendse F. & Snoo G.R. de (2016), Landscape complexity and farmland biodiversity: evaluating the CAP target on natural elements, Journal for Nature Conservation 30: 19-26.
- Speksnijder L., Oom D.M.J., Koning A.H.J., Biesmeijer C.S., Steegers E.A.P. & Steensma A.B. (2016), Agreement and reliability of pelvic floor measurements during rest and on maximum Valsalva maneuver using three-dimensional translabial ultrasound and virtual reality imaging, Ultrasound in Obstetrics and Gynecology 48(2): 243-249.
- Garratt M.P.D., Breeze T.D., Boreux V., Fountain M.T., McKerchar M., Webber S.M., Coston D.J., Jenner N., Dean R., Westbury D.B., Biesmeijer J.C. & Potts S.G. (2016), Apple pollination: demand depends on variety and supply depends on pollinator identity, PLoS ONE 11(5): e0153889.
- Aguirre-Gutierrez J., Kissling W.D., Carvalheiro L.G., WallisDeVries M.F., Franzen M. & Biesmeijer J.C. (2016), Functional traits help to explain half-century long shifts in pollinator distributions, Scientific Reports 6: 24451.
- Potts S.G., Imperatriz-Fonseca V., Ngo H.T., Aizen M.A., Biesmeijer J.C., Breeze T.D., Dicks L.V., Garibaldi L.A. Hill R., Settele J. & Vanbergen A.J. (2016), Safeguarding pollinators and their values to human well-being, Nature 540: 220-229.
- Potts S.G., Imperatriz-Fonseca V., Ngo H.T., Biesmeijer J.C., Breeze T.D., Dicks L.V., Garibaldi L.A., Hill R., Settele J. & Vanbergen A.J. (2016), Summary for policymakers of the thematic assessment on pollinators, pollination and food production. REVISTA BIOTA NEOTROPICA. [other].
- Kleijn D., Winfree R., Bartomeus I., Carvalheiro L.G., Henry M., Isaacs R., Klein A.M., Kremen C., M'Gonigle L.K., Rader R., Ricketts T.H., Williams N.M., Adamson N.L., Ascher J.S., Baldi A., Batary P., Benjamin F., Biesmeijer J.C., Blitzer E.J., Bommarco R., Brand M.R., Bretagnolle V., Button L., Cariveau D.P., Chifflet R., Colville J.F., Danforth B.N., Elle E., Garratt M.P.D., Herzog F., Holzschuh A., Howlett B.G., Jauker F., Jha S., Knop E., Krewenka K.M., Feon V. le, Mandelik Y., May E.A., Park M.G., Pisanty G., Reemer M., Riedinger V., Rollin O., Rundlof M., Sardinas H.S., Scheper J., Sciligo A.R., Smith H.G., Steffan-Dewenter I., Thorp R., Tscharntke T., Verhulst J., Viana B.F., Vaissiere B.E., Veldtman R., Westphal C. & Potts S.G. (2015), Delivery of crop pollination services is an insufficient argument for wild pollinator conservation, Nature Communications 6: 7414.
- Gerard M., Michez D., Fournier D., Maebe K., Smagghe G., Biesmeijer J.C. & Meulemeester T. de (2015), Discrimination of haploid and diploid males of Bombus terrestris (Hymenoptera; Apidae) based on wing shape, Apidologie 46: 644-653.
- Eskildsen A., Carvalheiro L.G., Kissling W.D., Biesmeijer J.C., Schweiger O. & Hoye T.T. (2015), Ecological specialization matters: long-term trends in butterfly species richness and assemblage composition depend on multiple functional traits, Diversity and Distributions 21(7): 792-802.
- Maebe K., Meeus I., Ganne M., Meulemeester T. de, Biesmeijer K. & Smagghe G. (2015), Microsatellite analysis of museum specimens reveals historical differences in genetic diversity between declining and more stable Bombus species, PLoS ONE 10: e0127870.
- Varma V., Ratnam J., Viswanathan V., Osuri A.M., Biesmeijer J.C., Madhusudan M.D., Sankaran M., Krishnadas M., Barua D., Budruk M., Isvaran K., Jayapal R., Joshi J., Karanth K.K., Krishnaswamy J., Kumar R., Mukherjee S., Nagendra H., Niphadkar M., Owen N., Page N., Prasad S., Quader S., Nandini R., Robin V.V., Sait S.M., Shah M.A., Somanathan H., Srinivasan U. & Sundaram B. (2015), Perceptions of priority issues in the conservation of biodiversity and ecosystems in India, Biological Conservation 187: 201-211.
- Senapathi D., Biesmeijer J.C., Breeze T.D., Kleijn D., Potts S.G. & Carvalheiro L.G. (2015), Pollinator conservation: the difference between managing for pollination services and preserving pollinator diversity, Current Opinion in Insect Science 12: 93-101.
- Ferreira P.A., Boscolo D., Carvalheiro L.G., Biesmeijer J.C., Rocha P.L.B. & Viana B.F. (2015), Responses of bees to habitat loss in fragmented landscapes of Brazilian Atlantic Rainforest, Landscape Ecology 30: 2067-2078.
- Aguirre-Gutierrez J., Biesmeijer J.C., Loon E.E. van, Reemer M., WallisDeVries M.F. & Carvalheiro L.G. (2015), Susceptibility of pollinators to ongoing landscape changes depends on landscape history, Diversity and Distributions 21(1): 1129-1140.
- Marshall L., Carvalheiro L.G., Aguirre-Gutierrez J., Bos M., Groot G.A. de, Kleijn D., Potts S.G., Reemer M., Roberts S., Scheper J. & Biesmeijer J.C. (2015), Testing projected wild bee distributions in agricultural habitats: predictive power depends on species traits and habitat type, Ecology and Evolution 5(19): 4426-4436.
- Senapathi D., Carvalheiro L.G., Biesmeijer J.C., Dodson C.A., Evans R.L., McKerchar M., Morton R.D., Moss E.D., Roberts S.P.M., Kunin W.E. & Potts S.G. (2015), The impact of over 80 years of land cover changes on bee and wasp pollinator communities in England, Proceedings of the Royal Society B: Biological Sciences 282(1806): 20150294.
- Breeze T.D., Vaissiere B.E., Bommarco R., Petanidou T., Seraphides N., Kozak L., Scheper J., Biesmeijer J.C., Kleijn D., Gyldenkaerne S., Moretti M., Holzschuh A., Steffan-Dewenter I., Stout J.C., Partel M., Zobel M. & Potts S.G. (2014), Agricultural policies exacerbate honeybee pollination service supply-demand mismatches across Europe, PLoS ONE 9(1): e82996.
- Garratt M.P.D., Breeze T.D., Jenner N., Polce C., Biesmeijer J.C. & Potts S.G. (2014), Avoiding a bad apple: insect pollination enhances fruit quality and economic value, Agriculture, Ecosystems and Environment 184: 34-40.
- Polce C., Garratt M.P., Termansen M., Ramirez-Villegas J., Challinor A.J., Lappage M.G., Boatman N.D., Crowe A., Endalew A.M., Potts S.G., Somerwill K.E. & Biesmeijer J.C. (2014), Climate-driven spatial mismatches between British orchards and their pollinators: increased risks of pollination deficits, Global Change Biology 20(9): 2815-2828.
- Thompson C.E., Biesmeijer J.C., Allnutt T.R., Pietravalle S. & Budge G.E. (2014), Parasite pressures on feral honey bees (Apis mellifera sp.), PLoS ONE 9(8): e105164.
- Sandrock C., Tanadini L.G., Pettis J.S., Biesmeijer J.C., Potts S.G. & Neumann P. (2014), Sublethal neonicotinoid insecticide exposure reduces solitary bee reproductive success, Agricultural and Forest Meteorology 16(2): 119-128.
- Elbgami T., Kunin W.E., Hughes W.O.H. & Biesmeijer J.C. (2014), The effect of proximity to a honeybee apiary on bumblebee colony fitness, development, and performance, Apidologie 45: 504-513.
- Garratt M.P.D., Coston D.J., Truslove C.L., Lappage M.G., Polce C., Dean R., Biesmeijer J.C. & Potts S.G. (2014), The identity of crop pollinators helps target conservation for improved ecosystem services, Biological Conservation 169: 128-135.
- Carvalheiro L.G., Biesmeijer J.C., Benadi G., Frund J., Stang M., Bartomeus I., Kaiser-Bunbury C.N., Baude M., Fernandes Gomes S.I., Merckx V.S.F.T., Baldock K.C.R., Bennett A.T.D., Boada R., Bommarco R., Cartar R., Chacoff N., Danhardt J., Dicks L.V., Dormann C.F., Ekroos J., Henson K.S.E., Holzschuh A., Junker R.R., Lopezaraiza-Mikel M., Memmott J., Montero-Castano A., Nelson I.L., Petanidou T., Power E.F., Rundlof M., Smith H.G., Stout J.C., Temitope K., Tscharntke T., Tscheulin T., Vila M. & Kunin W.E. (2014), The potential for indirect effects between co-flowering plants via shared pollinators depends on resource abundance, accessibility and relatedness, Ecology Letters 17(11): 1389-1399.
- Gonzalez-Varo J.P., Biesmeijer J.C., Bommarco R., Potts S.G., Schweiger O., Smith H.G., Steffan-Dewenter I., Szentgyorgyi H., Woyciechowski M. & Vila M. (2013), Combined effects of global change pressures on animal-mediated pollination, Trends in Ecology & Evolution 28(9): 524-530.
- Shackelford G., Steward P.R., Benton T.G., Kunin W.E., Potts S.G., Biesmeijer J.C. & Sait S.M. (2013), Comparison of pollinators and natural enemies: a meta-analysis of landscape and local effects on abundance and richness in crops, Biological Reviews 88(4): 1002-1021.
- Aguirre-Gutierrez J., Carvalheiro L.G., Polce C., Loon E.E. van, Raes N., Reemer M. & Biesmeijer J.C. (2013), Fit-for-purpose: species distribution model performance depends on evaluation criteria – Dutch hoverflies as a case study, PLoS ONE 8: e63708.
- Dicks L.V., Abrahams A., Atkinson J., Biesmeijer J., Bourn N., Brown C., Brown M.J.F., Carvell C., Connolly C., Cresswell J.E., Croft P., Darvill B., Zylva P. de, Effingham P., Fountain M., Goggin A., Harding D., Harding T., Hartfield C., Heard M.S., Heathcote R., Heaver D., Holland J., Howe M., Hughes B., Huxley T., Kunin W.E., Little J., Mason C., Memmott J., Osborne J., Pankhurst T., Paxton R.J., Pocock M.J.O., Potts S.G., Power E.F., Raine N.E., Ranelagh E., Roberts S., Saunders R., Smith K., Smith R.M., Sutton P., Tilley L.A.N., Tinsley A., Tonhasca A., Vanbergen A.J., Webster S., Wilson A. & Sutherland W.J. (2013), Identifying key knowledge needs for evidence-based conservation of wild insect pollinators: a collaborative cross-sectoral exercise, Insect Conservation and Diversity 6(3): 435-446.
- Giannini T.C., Chapman D.S., Saraiva A.M., Alves-dos-Santos I. & Biesmeijer J.C. (2013), Improving species distribution models using biotic interactions: a case study of parasites, pollinators and plants, Ecography 36(6): 649-656.
- Polce C., Termansen M., Aguirre-Gutierrez J., Boatman N.D., Budge G.E., Crowe A., Garratt M.P., Pietravalle S., Potts S.G., Ramirez J.A., Somerwill K.E. & Biesmeijer J.C. (2013), Species distribution models for crop pollination: a modelling framework applied to Great Britain, PLoS ONE 8(10): e76308.
- Carvalheiro L.G., Kunin W.E., Keil P., Aguirre-Gutierrez J., Ellis W.N., Fox R., Groom Q., Hennekens S., Landuyt W. van, Maes D., Meutter F. van de, Michez D., Rasmont P., Ode B., Potts S.G., Reemer M., Roberts S.P.M., Schaminee J., WallisDeVries M.F. & Biesmeijer J.C. (2013), Species richness declines and biotic homogenisation have slowed down for NW-European pollinators and plants, Ecology Letters 16(7): 870-878.
- Hendriks R.J.J., Carvalheiro L.G., Kleukers R.M.J.C. & Biesmeijer J.C. (2013), Temporal-spatial dynamics in Orthoptera in relation to nutrient availability and plant species richness, PLoS ONE 8: e71736.
- Marini L., Quaranta M., Fontana P., Biesmeijer J.C. & Bommarco R. (2012), Landscape context and elevation affect pollinator communities in intensive apple orchards, Basic and Applied Ecology 13(8): 681-689.
- Evison S.E.F., Roberts K.E., Laurenson L., Pietravalle S., Hui J., Biesmeijer J.C., Smith J.E., Budge G. & Hughes W.O.H. (2012), Pervasiveness of parasites in pollinators, PLoS ONE 7(1): e30641.
- Potts S.G., Biesmeijer J.C., Bommarco R., Felicioli A., Fischer M., Jokinen P., Kleijn D., Klein A.M., Kunin W.E., Neumann P., Penev L.D., Petanidou T., Rasmont P., Roberts S.P.M., Smith H.G., Sorensen P.B., Steffan-Dewenter I., Vaissiere B.E., Vila M., Vujic A., Woyciechowski M., Zobel M., Settele J. & Schweiger O. (2011), Developing European conservation and mitigation tools for pollination services: approaches of the STEP (Status and Trends of European Pollinators) project, Journal of Apicultural Research 50(2): 152-164.